0:00
introduction to clidamonus a fascinating
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genus of unicellular green
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algae these single-sellled organisms are
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found in various environments they
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thrive in freshwater ponds and lakes
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moist soil environments and even on damp
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tree bark chidamonus has two distinctive
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features that make it remarkable first
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chidamonus cells have two fleella which
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are whip-like structures used for
0:28
second they contain chloroplasts which
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allow them to photosynthesize converting
0:38
energy clomidamonus serves as an
0:40
important model organism in biological
0:42
research scientists use it to study
0:45
photosynthesis fleeller movement and
0:47
structure cellular responses to light
0:50
and for various genetic research and
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biotechnology applications to summarize
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cladomonus is a unicellular green algae
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with two fleella for movement and
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photosynthetic capability found in
1:05
environments now we'll examine the
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external features of clidamonus a
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fascinating unicellular green alga
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unlike plant cells that have cellulose
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cell walls clidamonus has a cell wall
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made primarily of glyoproteins arranged
1:21
a distinctive feature of clamidamonus is
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its two anterior fugella which emerge
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from the apical end of the
1:28
cell these fugella move in a coordinated
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breast stroke pattern during the power
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stroke the fleella sweep backward to
1:36
forward in the recovery stroke the
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fleella curl and return to their initial
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position with minimal resistance through
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this coordinated fleeller movement
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enables clidamonus to navigate through
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water toward light or
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nutrients to summarize clamdomonus has a
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glyoprotein cell wall and two interior
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fugella that enable its distinctive
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movement pattern for finding optimal
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nutrients clidamonus contains several
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specialized internal organels that
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perform essential cellular functions
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the most prominent organel is the
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cupshaped chloroplast it contains
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chlorophyll which captures light energy
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photosynthesis the chloroplast contains
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stacked membranes called philyloids
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where the light dependent reactions
2:31
occur within the chloroplast is a
2:34
specialized structure called the pyonoid
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the pyonoid is crucial for carbon
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fixation containing high concentrations
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of the enzyme rubiscoco which captures
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carbon dioxide for the Calvin
2:47
cycle the nucleus contains the cell's
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genetic material in the form of DNA it
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directs cellular activities by
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controlling gene expression within the
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nucleus is a darker region called the
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nucleolus where ribosomal RNA is
3:04
mitochondria are the powerhouses of the
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cell even though clamdomonus can
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photosynthesize it still needs
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mitochondria to generate ATP through
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cellular respiration especially when
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unavailable the Golgi apparatus
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processes and packages proteins and
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lipids produced in the cell these
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materials are then transported to other
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parts of the cell or secreted outside
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via small membranebound vesicles
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contractile vacules are specialized
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organels that regulate water balance
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since chidamonus often lives in
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freshwater environments water constantly
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enters the cell by osmosis these vacules
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collect excess water and periodically
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contract to expel it from the
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here we can see a complete
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cross-sectional view of a clidamonus
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cell showing the spatial arrangement of
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all internal organels note how the
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cup-shaped chloroplast occupies a large
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portion of the cell with the pyonoid
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embedded within it the nucleus is
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positioned toward the center while other
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organels like mitochondria golgi
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apparatus and contractile vacules are
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distributed throughout the cytoplasm to
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summarize each organel performs
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specialized functions that contribute to
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the overall survival and reproduction of
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chamomonus the chloroplast enables
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photosynthesis while the pyonoid
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enhances carbon fixation
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efficiency the nucleus directs cellular
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activities mitochondria provide energy
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the Golgi apparatus processes proteins
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and contractile vacules maintain water
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balance in freshwater environments
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clidamona species are incredibly
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adaptable microorganisms found in
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diverse habitats across the
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globe freshwater environments are the
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primary habitats for most clidimonus
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species they thrive in lakes ponds and
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slowmoving streams they typically prefer
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neutral to slightly alkaline pH and
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moderate temperatures between 20 and 25°
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their photosynthetic nature means
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they're commonly found in well-lit
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waters several chlamidona species have
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adapted to life in moist soil
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environments they're typically found in
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the top layers of nutrient-rich soils
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during dry periods these soil dwelling
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species can form protective cysts
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allowing them to remain dormant for
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extended periods until favorable
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remarkably certain chamomidonus species
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have adapted to extreme environments
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including snow and alpine regions
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clidamonus nalis is famous for creating
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the red snow phenomenon it produces
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carotenoid pigments that protect against
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intense UV radiation and help absorb
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heat these adaptations allow survival in
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freezing temperatures
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while most clidamonus species prefer
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fresh water several have adapted to
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brackish and even saline environments
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these salt tolerant species have evolved
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specialized ion transporters and
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mechanisms to regulate osmotic pressure
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they can be found in coastal waters salt
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marshes and estuaries with some species
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like chlidamonus pulsatilla thriving in
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fully marine environments
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clidamona species have a truly global
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distribution found on every continent
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including Antarctica they show
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remarkable adaptability across diverse
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ecosystems while greatest species
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diversity is observed in temperate
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regions specialized species have
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colonized extreme environments from sea
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level to alpine regions above 4,000 m
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clidamonus reproduces asexually by
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forming multiple daughter cells within
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wall the process begins with a mature
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clidamonus cell that prepares for
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division the asexual reproduction
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process consists of several key steps
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first the cell retracts its
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fleella next multiple nuclear divisions
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occur within the parent cell wall the
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cytoplasm then divides as separate cells
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begin to form finally multiple daughter
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cells form within the parent cell
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wall clamonus produces three main types
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of asexual spores under different
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environmental conditions zuspores are
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motile daughter cells with fleella
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formed under favorable conditions
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planospores are non-motile spores that
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develop when nutrients are limited
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hypnospores have thick protective walls
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and form during harsh environmental
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conditions different environmental
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factors trigger specific asexual
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reproductive strategies in climatonus in
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nutrient-rich water rapid cell division
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produces 2 to eight zoo spores when
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nitrogen is limited cells form
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non-motile alanoposopores during drought
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or cold conditions cells create
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resistant hypnospores with thick walls
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light and dark cycles can trigger
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synchronized division resulting in
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spores the asexual cell cycle of
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clamdomonus consists of distinct phases
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unlike most cells that divide by binary
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fision clidamonus underos multiple
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fision during its cell cycle growth
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occurs in G1 phase followed by DNA
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synthesis in Sphase after G2 preparation
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the cell undergoes multiple fishision in
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Mphase producing two to eight daughter
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cells at once unlike most ukarotes
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clamdomonus can undergo multiple rounds
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of DNA replication and nuclear division
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divides sexual reproduction in
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clidamonus is triggered by environmental
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stress factors these stressors include
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nitrogen depletion temperature changes
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and fluctuations in light
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intensity clidamonus exhibits three
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distinct types of sexual reproduction
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isogamy occurs when two gametes of
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similar size and appearance fuse
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together anogamy involves gameamtes of
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different sizes though both remain
9:24
ugami features a large non-motile egg
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cell and a small motile sperm
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cell after fusion the resulting zygote
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underos a series of developmental stages
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the process begins with the formation of
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a diploid zygote after gameamt fusion
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the zygote develops a thick protective
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wall becoming a zygospor that can
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survive harsh conditions the zygospore
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enters a dormant phase which can last
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for extended periods during unfavorable
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conditions when conditions improve
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meiosis occurs within the zygospor
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producing hloid cells the zygospor wall
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ruptures releasing four hloid cells that
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can grow into new clidamonus
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individuals to summarize the sexual
10:08
reproduction cycle in clamidonus begins
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with environmental stress which triggers
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gameamt formation these gametes then
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fuse forming a zygote that develops into
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a dormant zygospore when conditions
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improve meiosis occurs completing the
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cycle by releasing hloid cells that grow
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into new individuals clidamonus has a
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tripartite genome structure meaning its
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genetic material is distributed across
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three distinct cellular compartments
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these three locations are the nucleus
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which is the main genetic control center
10:40
the chloroplast which enables
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photosynthesis and the mitochondrian
10:46
energy the nuclear genome of clamdomonus
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consists of 17 chromosomes with
10:52
approximately 120 million base pairs it
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contains 15,000 genes
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the chloroplast genome has a distinct
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circular DNA structure consisting of
11:03
about 200,000 base pairs and encoding
11:06
approximately 100 genes it's maternally
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inherited and contains genes essential
11:13
photosynthesis unlike most mitochondrial
11:16
genomes the mitochondrial DNA of
11:18
chlidamonus has a linear structure it's
11:21
relatively small with only about 15 to
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20,000 base pairs encoding approximately
11:27
20 genes related to cellular
11:29
respiration the tripartite genome
11:32
structure of clamonus makes it
11:34
exceptionally valuable for research it
11:36
serves as an ideal model for studying
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organel genome evolution and
11:40
understanding the coordination between
11:42
nuclear and organel genes clidamonus is
11:46
unique among model organisms because all
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three of its genomes are accessible for
11:50
experimental manipulation enabling
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researchers to study genetic
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interactions across different cellular
11:58
compartments the three genomes differ
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dramatically in size the nuclear genome
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is the largest at about 120 million base
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pairs while the chloroplast genome is
12:08
much smaller at around 200,000 base
12:10
pairs the mitochondrial genome is the
12:14
smallest with only about 20,000 base
12:17
pairs together these three genomes
12:20
contain all the genetic information
12:22
needed for clamonus to grow reproduce
12:26
environment the genus clamonus includes
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over 600 species with diverse
12:31
adaptations to varied environments
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including fresh water soil and even snow
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clidamonus rhinardi is the primary model
12:40
organism used in research it has a fully
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sequenced 120 megabase genome can be
12:46
easily cultured in laboratories and its
12:48
hloid genetics simplifies genetic
12:51
analysis clidamonus nalis is remarkable
12:54
for containing red carotenoid pigments
12:56
that protect it from intense ultraviolet
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radiation at high altitudes this species
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thrives in alpine and polar snow fields
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causing the phenomenon known as
13:06
watermelon snow pink or red colored snow
13:09
patches beyond the well-known species
13:12
chidamonus includes many others with
13:14
remarkable adaptations clidamonus
13:17
acidophila thrives in highly acidic
13:19
waters with specialized cell walls
13:21
clomidamonus uriel has adapted to marine
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environments with salt tolerance
13:27
clidamonus musi is found in soil and
13:30
fresh water with efficient nutrient
13:32
uptake strategies and chidamonus
13:34
debariana survives in temporary water
13:36
bodies through rapid reproduction cycles
13:41
zygot clidamona species have evolved
13:44
remarkable adaptations to thrive in
13:46
diverse environments their temperature
13:48
tolerance ranges fromus35 to plus 35°
13:53
different species have adapted to pH
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levels from highly acidic to alkaline
13:58
conditions they show impressive light
14:00
adaptations from high UV resistance to
14:03
efficient photosynthesis in low light
14:06
their nutrient requirements vary widely
14:08
allowing them to colonize environments
14:10
from pristine snow fields to
14:12
nutrient-rich soils and waters these
14:15
diverse adaptations across chidamonus
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species make them not only fascinating
14:19
research subjects but also important
14:21
components of various ecosystems
14:24
worldwide clidamonus Reinhardy serves as
14:27
an exceptional model organism in
14:29
biological research its rapid growth
14:32
cycle hloid genetics transformable
14:34
genome and fully sequenced DNA make it
14:37
an ideal subject for laboratory studies
14:39
researchers utilize clamdomonus to
14:41
investigate fundamental processes like
14:43
photosynthesis fleeller movement cell
14:46
cycle regulation and chloroplast
14:48
development in natural ecosystems
14:51
clamadomonus serves as a primary
14:52
producer generating oxygen sequestering
14:55
carbon and providing a crucial food
14:57
source for aquatic organisms beyond
15:00
basic research clamdomonus has practical
15:03
applications in bofuel production
15:05
recombinant protein expression
15:07
environmental remediation and water
15:09
quality assessment current research
15:11
continues to expand clatomonus
15:13
applications with crisper gene editing
15:16
climate change studies synthetic biology
15:18
approaches and improved bofuel
15:20
production technologies
15:22
the significance of Clamatomonus
15:24
continues to grow as new technologies
15:26
enhance our ability to study and utilize
15:29
this remarkable model organism